The story begins in Feudal Japan, when the half-demon InuYasha steals the Jewel of Four Souls—a magical jewel that enhances the power of and can grant a single wish to the one who wields it—from a village. InuYasha desires the gem in order to turn himself into a full blooded demon. InuYasha does not get far before Kikyo, the young miko of the village, shoots him with a sacred arrow that indefinitely seals him onto Goshinboku, a sacred tree in the nearby forest. After being mortally wounded, Kikyo tells her younger sister, Kaede, to burn the jewel with her body to prevent it from falling into the hands of evil.
In modern Tokyo, a junior high school girl named Kagome Higurashi is on her way to school. She stops in the well house of her family's Shinto shrine to retrieve her cat, Buyo, when a centipede demon emerges from the well and pulls her into the Sengoku period of Japan.
Not knowing where she is, Kagome wanders around a forest near the Bone Eater's Well. She spots Goshinboku off in the distance and proceeds towards it. On the tree, she finds InuYasha, who is still sealed in an enchanted sleep. Villagers seize her and take her to the old priestess, Kaede. Recognizing that Kagome is the reincarnation of her sister Kikyo, Kaede tells her the story of Kikyo and InuYasha.
The centipede attacks again, and Kagome is forced to release InuYasha from his enchantment so he can kill it. After defeating the centipede, InuYasha tries to take the Jewel of Four Souls from Kagome. In order to thwart InuYasha and to save Kagome's life, Kaede places magical prayer beads around InuYasha's neck so that Kagome can subdue him.
The Jewel of Four Souls attracts more demons, and in a battle against a carrion crow demon, the jewel is shattered into numerous shards that spread all across Japan. As the individual shards are capable of granting those who embed it in their bodies with great power, it is sought after by many demons as well as some humans.
Kagome and InuYasha set out to collect the shards and restore the Jewel of Four Souls. Along the way, they befriend Shippo, a fox demon; Miroku, a cursed monk; and Sango, a demon-slayer whose brother was possessed and forced to slaughter their father and the other demon slayers. The group encounters many friends and foes during the adventure, including InuYasha's older half-brother Sesshomaru; Kikyo, who died 50 years ago but, was resurrected with part of Kagome's soul; Naraku, who tricked Kikyo and InuYasha into turning against each other; and a wolf demon named Koga, who is in love with Kagome and whenever he and InuYasha meet they constantly bicker.
Eventually, the Shikon jewel is reassembled by Naraku, who is then defeated by InuYasha. Before he dies, Naraku makes a wish on the jewel that Kagome would replace the soul of Midoriko, the jewel's original creator, within the jewel and that his soul would also enter the jewel so he could survive inside as her enemy. This would only work if Kagome made a selfish wish, which Naraku tries to trick her into doing, but confident that InuYasha would rescue her, she refuses to do so. InuYasha rescues her and Kagome wishes for the jewel to disappear forever.
Kagome is thrown back into her own time, and InuYasha is no longer able to see her. However, after three years when Kagome has graduated high school, a portal opens allowing her to return to InuYasha's time. They acknowledge their love for one another and she chooses to remain in the past with him.
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